Do you have the "Air intake control button" labeled "9" on page 279 of the 2010 RX350 navigation manual? If you have this button, pressing it repeatedly cycles the system through three modes -- exactly the same as a similar button works in my 2000 LS400. The three modes are described on page 284 of the 2010 RX350 navigation manual.
I will include page 303 from the 2010 RX350 navigation manual simply because it points to page 410 which (poorly) shows where the sensitivity is customized by selecting "Vehicle Customization" and then "Climate Settings".
Have you gone into the Climate Settings screen to see if has the sensitivity customization feature?
If your RX350 isn't like what is shown on the attached 2010 RX350 navigation manual pages, could you take a photo of the buttons you have and attach it to this thread?
If your A/C system is like what is shown on the attached pages, then you have a system that automatically adjusts between outside air and recirculation depending on outside air quality.
Your experience with Lexus dealership staff not knowing much about the cars they sell certainly matches what I have experience over the past 20 years. My "favorite" experience was trying to convince two service writers that my 00 LS400 has DRL. They argued forcefully that DRL didn't start until the 2001 model year. They didn't even apologize when I showed them information about DRL in my owners manual.
Thanks for the photos--I will take a photo of mine, but in my manual they show an 'A' version and a 'B' version. One version looks like the photos you've attached, however the other is what I have in my car--where the word 'Auto' appears on the button in your photos, I have a second car icon with a circulation arrow in it--just like most cars with internal/external air circulation. When I look on the Lexus website, it appears that at some point (recently), they moved the auto feature into the luxury package. Earlier documentation shows this was part of the Nav. I tried to get the luxury package, but they don't seem to exist in New England....Thanks for your help with this!